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The wolf Canis lupus , [a] also known as the gray wolf , timber wolf , [4] [5] western wolf , [b] and its other subspecies is a canine native to the wilderness and remote areas of Eurasia and North America.

The gray wolf is the second most specialised member of the genus Canis , after the Ethiopian wolf , as demonstrated by its morphological adaptations to hunting large prey, its more gregarious nature, [10] and its highly advanced expressive behavior. It is the only species of Canis to have a range encompassing both the Old and New Worlds , [6] and originated in Eurasia during the Pleistocene , colonizing North America on at least three separate occasions during the Rancholabrean.

It feeds primarily on large ungulates , though it also eats smaller animals, livestock, carrion, and garbage. The global gray wolf population is estimated to be , Non-rabid wolves have attacked and killed people, mainly children, but this is rare, as wolves are relatively few, live away from people, and have developed a fear of humans from hunters and shepherds. The Latin lupus is a Sabine loanword.

The species Canis lupus was first recorded by Carl Linnaeus in his publication Systema Naturae in , [3] with the Latin classification translating into the English words "dog wolf". The thirty-seven subspecies of Canis lupus are listed under the designated common name of "wolf" in Mammal Species of the World third edition that was published in The evolution of the wolf occurred over a geologic time scale of at least thousand years.

The gray wolf Canis lupus is a highly adaptable species that is able to exist in a range of environments and which possesses a wide distribution across the Holarctic. Studies of modern gray wolves have identified distinct sub-populations that live in close proximity to each other. The archaeological and paleontological records show gray wolf continuous presence for at least the last , years. However, the geographic origin of this radiation is not known.

In , a genetic study found that the wolf population in Europe was divided along a north-south axis and formed five major clusters. Three clusters were identified occupying southern and central Europe in Italy, the Dinaric- Balkans , the Carpathians.

Another two clusters were identified occupying north-central Europe and the Ukrainian steppe. The Italian wolf consisted of an isolated population with low genetic diversity. Wolves from Finland, Latvia, Belarus, Poland and Russia formed the north-central Europe cluster, with wolves from the Carpathians cluster coming from a mixture of wolves from the north-central cluster and the Dinaric-Balkans cluster.

The wolves from the Carpathians were more similar to the wolves from the Pontic-Caspian Steppe than they were to wolves from north-central Europe.

These clusters may have been the result of expansion from glacial refugia , an adaptation to local environments, and landscape fragmentation and the killing of wolves in some areas by humans.

In , two genetic studies of North American gray wolves found that they formed six ecotypes — genetically and ecologically distinct populations separated from other populations by their different type of habitat. The studies found that precipitation and mean diurnal temperature range were the most influential variables. It was once thought that dogs and gray wolves did not voluntarily interbreed in the wild, though they can produce fertile wolf-dog offspring.

The gray wolf has interbred extensively with the eastern wolf producing a hybrid population termed Great Lakes boreal wolves. Gray wolf Y-chromosomes have also been found in Texan coyote haplotypes. At six months of age, the hybrids were closely monitored and were shown to display both physical and behavioral characteristics from both species.

The gray wolf is the largest extant member of the Canidae , excepting certain large breeds of domestic dog. Compared to its closest wild cousins the coyote and golden jackal , the gray wolf is larger and heavier, with a broader snout, shorter ears, a shorter torso and longer tail.

The gray wolf usually carries its head at the same level as the back, raising it only when alert. Generally, wolves have a high heart weight of 0. Tibetan gray wolves, which occupy territories up to 3, above sea level , have evolved hearts that withstand the low oxygen levels.

The gray wolf's head is large and heavy, with a wide forehead, strong jaws and a long, blunt muzzle. This force is sufficient to break open most bones. A similar trend was found with the carnassial tooth bite force, but with the extinct dire wolf and gray wolf both measuring , then followed by the African hunting dog , the dhole , and the dingo The gray wolf has very dense and fluffy winter fur, with short underfur and long, coarse guard hairs.

Especially long hairs are on the shoulders, and almost form a crest on the upper part of the neck. The hairs on the cheeks are elongated and form tufts.

The ears are covered in short hairs, which strongly project from the fur. Short, elastic and closely adjacent hairs are present on the limbs from the elbows down to the calcaneal tendons. Wolf fur provides better insulation than dog fur, and does not collect ice when warm breath is condensed against it. Older wolves generally have more white hairs in the tip of the tail, along the nose and on the forehead.

The winter fur is retained longest in lactating females, though with some hair loss around their nipples. Coat color ranges from almost pure white through various shades of blond, cream, and ochre to grays, browns, and blacks, [68] with variation in fur color tending to increase in higher latitudes. The gray wolf is a social animal, whose basic social unit consists of a mated pair, accompanied by the pair's adult offspring. In the rare cases where other wolves are adopted, the adoptee is almost invariably an immature animal 1—3 years of age unlikely to compete for breeding rights with the mated pair.

In some cases, a lone wolf is adopted into a pack to replace a deceased breeder. Wolves are highly territorial animals, and generally establish territories far larger than they require to survive in order to assure a steady supply of prey.

Territory size depends largely on the amount of prey available and the age of the pack's pups, tending to increase in size in areas with low prey populations [77] or when the pups reach the age of 6 months, thus having the same nutritional needs as adults.

Wolves defend their territories from other packs through a combination of scent marking , direct attacks and howling see Communication. Scent marking is used for territorial advertisement, and involves urination, defecation and ground scratching. Such markers can last for 2—3 weeks, [78] and are typically placed near rocks, boulders, trees, or the skeletons of large animals.

The gray wolf is generally monogamous , [86] with mated pairs usually remaining together for life. Upon the death of one mated wolf, pairs are quickly re-established. Since males often predominate in any given wolf population, unpaired females are a rarity. Such gray wolves are termed " Casanova wolves" and, unlike males from established packs, they do not form pair bonds with the females they mate with.

Some gray wolf packs may have multiple breeding females this way, as is the case in Yellowstone National Park. This might take place if the original parents die or are for some reason separated from them. The age of first breeding in gray wolves depends largely on environmental factors: This is further demonstrated by the fact that captive wolves have been known to breed as soon as they reach 9—10 months, while the youngest recorded breeding wolves in the wild were 2 years old.

Females are capable of producing pups every year, with one litter annually being the average. Unlike the coyote, the gray wolf never reaches reproductive senescence. The gestation period lasts 62—75 days, with pups usually being born in the summer period.

Wolves bear relatively large pups in small litters compared to other canid species. The milk canines erupt after one month. Pups first leave the den after 3 weeks. Mother wolves do not leave the den for the first few weeks, relying on the fathers to provide food for them and their young. Pups begin to eat solid food at the age of 3—4 weeks. Pups have a fast growth rate during their first four months of life: Actual fights to establish hierarchy usually occur at 5—8 weeks of age.

This is in contrast to young foxes and coyotes, which may begin fighting even before the onset of play behavior. Although social animals, single wolves or mated pairs typically have higher success rates in hunting than do large packs, with single wolves having occasionally been observed to kill large prey such as moose , bison and muskoxen unaided. Because of this, it rarely manages to capture hidden hares or birds, though it can easily follow fresh tracks.

The actual killing method varies according to prey species. With large prey, mature wolves usually avoid attacking frontally, instead focusing on the rear and sides of the animal. Large prey, such as moose, is killed by biting large chunks of flesh from the soft perineum area, causing massive blood loss. With small, mouse -like prey, wolves leap in a high arc and immobilize it with their forepaws. Such instances are common in domestic animals, but rare in the wild.

In the wild, surplus killing primarily occurs during late winter or spring, when snow is unusually deep thus impeding the movements of prey [] or during the denning period, when wolves require a ready supply of meat when denbound. Once prey is brought down, wolves begin to feed excitedly, ripping and tugging at the carcass in all directions, and bolting down large chunks of it. When food is scarce, this is done at the expense of other family members, especially non-pups. Once the breeding pair has finished eating, the rest of the family tears off pieces of the carcass and transport them to secluded areas where they can eat in peace.

Wolves typically commence feeding by consuming the larger internal organs of their prey, such as the heart , liver , lungs and stomach lining. The kidneys and spleen are eaten once they are exposed, followed by the muscles. The gray wolf's expressive behavior is more complex than that of the coyote and golden jackal, as necessitated by its group living and hunting habits. While less gregarious canids generally possess simple repertoires of visual signals, wolves have more varied signals that subtly inter grade in intensity.

Passive submission usually occurs as a reaction to the approach of a dominant animal , and consists of the submissive wolf lying partly on its back and allowing the dominant wolf to sniff its anogenital area. Active submission occurs often as a form of greeting, and involves the submissive wolf approaching another in a low posture, and licking the other wolf's face.

The mouthing of each other's muzzles is a friendly gesture, while clamping on the muzzle with bared teeth is a dominance display. Similar to humans, gray wolves have facial color patterns in which the gaze direction can be easily identified, although this is often not the case in other canid species.

In , a study compared the facial color pattern across 25 canid species. The results suggested that the facial color pattern of canid species is related to their gaze communication, and that especially gray wolves use the gaze signal in conspecific communication. Gray wolves howl to assemble the pack usually before and after hunts , to pass on an alarm particularly at a den site , to locate each other during a storm or unfamiliar territory and to communicate across great distances.

Pups almost never howl, while yearling wolves produce howls ending in a series of dog-like yelps. The pitch usually remains constant or varies smoothly, and may change direction as many as four or five times. When pursuing prey, they emit a higher pitched howl, vibrating on two notes. When closing in on their prey, they emit a combination of a short bark and a howl.

The two are however mutually intelligible , as North American wolves have been recorded to respond to European-style howls made by biologists. Other vocalisations of wolves are usually divided into three categories:

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This article is about the gray wolf, canis lupus , and its subspecies. For other species of wolf and other uses, see Wolf disambiguation. For other uses, see Grey Wolf disambiguation. Middle Pleistocene — present , years BP [1]. Linnaeus , [3]. Subspecies of Canis lupus. Evolution of the wolf. List of gray wolf populations by country. Wolves in folklore, religion and mythology. Wolf attacks on humans and List of wolf attacks. Wolf hunting and Wolf hunting with dogs.

Human uses of hunted wolves. Wolves as pets and working animals. However, neither dogs nor dingoes are referred to as gray wolves. Throughout this article, the term "wolf" will be used to refer collectively to naturally occurring subspecies, especially the nominate subspecies, Canis lupus lupus. This terminology was first used in by Rudolf Schenkel of the University of Basel , who based his findings on researching the behavior of captive gray wolves.

This view on gray wolf pack dynamics was later popularized by L. David Mech in his book The Wolf. He formally disavowed this terminology in , explaining that it was heavily based on the behavior of captive packs consisting of unrelated individuals, an error reflecting the once prevailing view that wild pack formation occurred in winter among independent gray wolves.

Later research on wild gray wolves revealed that the pack is usually a family consisting of a breeding pair and its offspring of the previous 1—3 years. Bulletin of the American Museum of Natural History. International Union for Conservation of Nature. Tomus I in Latin 10 ed.

Retrieved November 23, Gray wolf Canis lupus and allies", in Feldhamer, George A. Wild Mammals of North America: Grey wolf Canis lupus Linnaeus, , pp. Foxes, Wolverhampton Wanderers F. Gland, Switzerland and Cambridge, UK. Mammal Species of the World: A Taxonomic and Geographic Reference 3rd ed. Johns Hopkins University Press.

Mammals of the Soviet Union Vol. Its Domestication and Behavior. The cautionary case of Bulgarian wolves". The Fear of Wolves: Norsk institutt for naturforskning NINA. A contribution to the study of wolf-dog hybridization patterns". Archived from the original PDF on September 24, Retrieved June 2, A new profile for the wolf. Journal of Vertebrate Paleontology. Walker's Mammals of the World. Electrocardiographic consequences of a peripatetic lifestyle in gray wolves Canis lupus.

Comparative Biochemistry and Physiology Part A: PLoS Genetics, 10 7. Comparative bite force in big biting mammals and the prediction of predatory behaviour in fossil taxa". Proceedings of the Royal Society B: Journal of Applied Genetics.

Canadian Journal of Zoology. Archived from the original on December 14, Environmental and intrinsic correlates of stress in free-ranging wolves. PLOS One, 10 9. Decade of the Wolf: The Quarterly Review of Biology. University of California Press. Animal Homosexuality and Natural Diversity. The Wolves of Isle Royale. Fauna of the National Parks of the United States. Ecology and conservation of wolves in a changing world. Retrieved March 16, Social behavior of langur monkeys Presbytis entellus in a high altitude environment.

Wolves Change An Ecosystem". Smithsonian National Zoological Park. Archived from the original on January 24, Retrieved August 10, Conservation Action Plan for the golden jackal Canis aureus L. S; Andriashek, D Mashkour "Hyenas around the cities. Zoology in the Middle East. Retrieved February 20, It's a Lot Like Cats and Dogs".

Mammals of the Soviet Union: Carnivora hyaenas and cats , Volume 2. Archived from the original on February 25, Wild cats of the world , University of Chicago Press, p. American Society of Mammalogists. Archived from the original PDF on June 17, In Large Carnivores and the Conservation of Biodiversity.

Carnivores quaternaires du Sud-Est de la France. In Weiss, Steven; Ferrand, Nuno. Phylogeography of Southern European Refugia. Archived from the original PDF on March 25, Also available in German: Heinz Staffelbach, Handbuch Schweizer Alpen. Animals in the Third Reich: The New York Times.

Retrieved September 4, Kuolleissa susissa vanhoja hauleja" , Iltalehti , March 19, Retrieved May 19, Translated by Kevin White. Retrieved May 2, Retrieved November 24, Mammals of the Holy Land. Texas Tech University Press, pp. Archived from the original PDF on August 21, Retrieved June 27, Cultural Perspectives , Psychology Press, pp. Archived from the original PDF on July 27, Smith, Yan Xie, Robert S. A Guide to the Mammals of China.

Retrieved February 9, Montana Department of Fish, Wildlife and Parks. Retrieved July 2, Archived from the original on July 30, Wisconsin Department of Natural Resources. Archived from the original PDF on March 19, Retrieved January 25, Snowmobile activity and glucocorticoid stress responses in wolves and elk.

Conservation Biology,16 3 , The Canadian Historical Review. Archived from the original PDF on June 21, A Complete Guide to Heraldry. University of Oklahoma Press. Oxford University Press, Oxford, pp.

The complex interactions between wolves Canis lupus and dogs Canis familiaris in a conservation context". Wolf and dog competition in Italy. Wolf attacks on dogs in Scandinavia — Will wolves in Scandinavia go extinct if dog owners are allowed to kill a wolf attacking a dog?

What's New for Conserving Carnivores". What is wrong with Romanian livestock guarding dogs? Hunting the grisly and other sketches; an account of the big game of the United States and its chase with horse, hound, and rifle , New York, London, G. Wolf and coyote trapping; an up-to-date wolf hunter's guide, giving the most successful methods of experienced "wolfers" for hunting and trapping these animals, also gives their habits in detail , Columbus, Ohio, A.

African palm civet N. Angolan slender mongoose G. Ethiopian dwarf mongoose H. Family Viverridae includes Civets. Small-toothed palm civet A. Sulawesi palm civet M. Masked palm civet P. Golden wet-zone palm civet P. Owston's palm civet C. Hose's palm civet D. Banded palm civet H.

Malabar large-spotted civet V. Small Indian civet V. Broad-striped Malagasy mongoose G. American black bear U. Molina's hog-nosed skunk C.

Sunda stink badger M. Southern spotted skunk S. Eastern lowland olingo B. Western mountain coati N. South American fur seal A. Northern fur seal C. Steller sea lion E. Australian sea lion N. South American sea lion O. New Zealand sea lion P. California sea lion Z.

Northern elephant seal M. Mediterranean monk seal M. Family Canidae includes dogs. African wild dog L. African clawless otter A. North American river otter L. Saharan striped polecat I. African striped weasel P. Hinny Lord Morton's mare Mule Zebroid.

Humanzee Hybrid orangutan Koolakamba. Polecat-ferret hybrid Polecat-mink hybrid. ABC Islands bear Grizzly—polar bear hybrid. Armiger Law of heraldic arms Grant of arms Blazon. Conventional elements of coats of arms Escutcheon. National coat of arms Arms of dominion Ecclesiastical heraldry Burgher arms Civic heraldry Canting arms Attributed arms.

Dolphin Ged Seahorse Lucy esox Scallop. Reremouse Bee Crapaudy toad Emmet ant Serpent. Phaleristics List of oldest heraldry Women in heraldry Socialist heraldry Vexillology.

Retrieved from " https: Uses authors parameter CS1 Latin-language sources la Articles with French-language external links Webarchive template wayback links Articles with Finnish-language external links CS1 maint: Views Read View source View history. In other projects Wikimedia Commons Wikispecies Wikiquote. This page was last edited on 28 October , at By using this site, you agree to the Terms of Use and Privacy Policy.

Eurasian wolf Canis lupus lupus. Canis lupus Linnaeus , [3]. Problems playing this file? Wikimedia Commons has media related to Canis lupus. Wikispecies has information related to Canis lupus. Atilax Marsh mongoose A. Crocuta Spotted hyena C. Large family listed below.

Small family listed below. Chrotogale Owston's palm civet C. Prionodon Banded linsang P. Civettictis African civet C. Galidia Ring-tailed mongoose G. Mature , Wife Added 3 day s ago From: Solo , Masturbating Added 1 month es ago From: Interracial , Cheating , Cuckold Added 5 month es ago From: Caged hubby waits for creampie Tags: Mature , French , Wife Added 4 month es ago From: Mature , Wife Added 6 year s ago From: Black on white scene of muscled bull and blonde Mature , Wife , Young Added 6 month es ago From: Mature , French , Wife Added 3 week s ago From: Interracial , Cheating , Cuckold Added 7 month es ago From: Interracial , Cuckold , Hd Added 1 month es ago From: Machines Added 4 month es ago From: Hot Sex Tube You Rate Tube Pretty Nu TV Nasty Video Tube Private Porn Films My Loved Video Full Red Sex Alex Pix Tube American, British, and French nationals are thought to be among the dead.

In other news, the latest figures from Angola indicate that at least , Chinese nationals have migrated to the country. The Angolan government says that work permits were issued for the Chinese to assist with development projects. To see the headlines and the articles, open the full news post. Thanks to Andy Bostom, C. Commenters are advised to leave their comments at this post rather than with the news articles so that they are more easily accessible. Gates of Vienna cannot vouch for the authenticity or accuracy of the contents of any individual item posted here.

We check each entry to make sure it is relatively interesting, not patently offensive, and at least superficially plausible.

Further research and verification are left to the reader. Vienna is the most fully enriched location, and seems to be in roughly the same situation as Oslo. Many thanks to Hermes for the translation from Unzensuriert. The number of pupils with a first language other than German has doubled from to The doubling in the number of multilingual students is evident in virtually all provinces in Austria.

The Styrian province is the only outlier, where the number of children with a different native language has tripled from about 5, to 16, The nationwide statistics conceals the dramatic developments in Vienna, because German is hardly spoken in the primary schools of the federal capital.

For a complete listing of previous enrichment news, see The Cultural Enrichment Archives. Scroll down for other posts that have appeared since Wednesday. Certain posts at Gates of Vienna, among them those by Takuan Seiyo, tend to attract the attention and comments of people who are preoccupied with the Jews. I generally delete such comments without publishing them. Before I deleted it, I sent it to Takuan, just to show him what was coming in.

He suggested that I go ahead and post it, followed by his response. Some interesting points, a lot of waffle and some errors.

Top class, modern historians now regard that figure as grotesquely disproportionate. It is one of the most successful internationsal businesses of all time.

You have the temerity to post this kind of Nazi apologist garbage in a comments thread of an article by a writer whose gentile grandparents had been murdered in a concentration camp and whose gentile mother spent two years in a labor-extermination camp, was a state witness in the post-war trial of its commandant, and relayed her wartime experiences to this writer directly.

Moreover, a writer who was born and grew up one hour by car from Auschwitz and three kilometers from the plant where the firm Hoch und Tiefbau AG had built the crematoria for that camp. In which, alone, 2. Moreover, you desecrate the memory of Witold Pilecki plus other Polish officer escapees from Auschwitz who produced written reports, e. I am omitting here reports by Jewish escapees, for example the Vrba-Wetzler report, as well as the fate of the Jewish part of my family during the war, so as to skirt the whole specious Joooos-tainted-it aspect of your comment.

The lowest for Auschwitz, for instance by the Polish historian Franciszek Piper , cites 1. The highest figure cited for Auschwitz is 4 million. However, much has happened since it went up, including the Blogger outage. Scroll down for a report on that. More new posts will be added below this one. The essay below is the conclusion of the ninth part in a series by Takuan Seiyo.

See the list at the bottom of this post for links to the previous installments. For over 60 years, White mea-culpists have had a firm grip in all fields of cultural mind imprinting: Their main endeavor has been to enforce their compulsory e. K and discretionary e. Nor the evils of the worldwide Islamic Inquisition which — not in the 16th century but now, in the 21st, condemns Muslim apostates to barbaric execution.

Instead, aggressive White androphobes of all genders which I can no longer count are decimating the philogynous and egalitarian West. Equality psychos are tearing down the most egalitarian society that ever existed except for initial communist experiments, before they turned bloody. American Jews, at the apex of the greatest fortune and philosemitic tolerance their long diaspora has ever bestowed on their kind, are busy supporting all the ideologies and policies that demolish their safe harbor and build up their Muslim, Black and Third World enemies.

Leftoid masochists and the Christian meek call for returning Hawaii to the Hawaiians and capitulating before a massive Mexican reconquista of one-third of America. The rightful Etruscan landowners are not bearing angry placards in front of the Vatican.

The Japanese are not planning to relinquish Hokkaido to its original owners, the Ainu. The tall, white and fair-haired Chachapoyas of the Andean forest have, alas, no remnants left to sue the Incas for genocide in a Peruvian court of law. However, even that great moral abyss of Western civilization — the Holocausts — stands out more in its industrialized and organizational features than it does either in the quality of its hatefulness or its relative or even absolute volumes.

In relative numbers, in just one year, , the Hutus and Tutsis in Rwanda, killed off a total of one million, in a population of 7 million.

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